Products Acknowledging Liverwort AToL Award(EF-0531750) and Abstracts.




  1. Long, D. G., Forrest, L. L., Villarreal, J. C. & Crandall-Stotler, B. J. 2016. Taxonomic changes in Marchantiaceae, Corsiniaceae and Cleveaceae (Marchantiidae, Marchantiophyta). Phytotaxa 252: 77-80.

  2. Söderström, L., Hagborg, A. , von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E. , Cargill, D. C., Costa, D. P., Crandall-Stotler, B. J. & 31 additional authors. 2016. World checklist of hornworts and liverworts. PhytoKeys 59: 1-828. (doi: 10.3897/phytokeys.59.6261). [This list of all recognized taxa of hornworts and liverworts was compiled by the first 3 authors, from contributions provided by the other authors involved in this very large project. All authors further edited and reviewed the manuscript as it was developed.]
  3. Long, D. G., L. L. Forrest, J. C. Villarreal & B. J. Crandall-Stotler. 2016. The genus Aitchisoniella Kashyap (Marchantiopsida, Cleveaceae) new to China, and its taxonomic placement. Journal of Bryology DOI: 10.1080/03736687.2016.1183072.



  1. Shaw, B., B. Crandall-Stotler, J. Vána, R. E. Stotler, M. von Konrat, J. J. Engel, E. C. Davis, D. G. Long, P. Sova & A. J. Shaw. 2015. Phylogenetic relationships and morphological evolution in a major clade of leafy liverworts (Phylum Marchantiophyta, Order Jungermanniales): Suborder Jungermanniineae. Systematic Botany 40: 27-45.

  2. Söderström, L., J. Vána, B. Crandall-Stotler, M. A. M. Renner, A. Hagborg & M. von Konrat. 2015. Notes on early land plants today. 68. Miscellaneous notes on Marchantiophyta. Phytotaxa 202: 69-72.

  3. Crandall-Stotler, B. 2015. From morphology to molecules: changing views of liverwort phylogeny and classification. Abstract for Botany 2015, Edmonton, Alberta Canada.

  4. Larráin, J., B. Carter, B. Shaw, J. Hentschel, L. S. Strozier, T. Furuki, J. Heinrichs, B. Crandall-Stotler, J. Engel & M. von Konrat. 2015. A resurrection of Neohattoria Kamim. (Jubulaceae, Marchantiophyta): a six decade systematic conflict resolved through a molecular perspective. PhytoKeys 50: 101-122.

  5. Söderström, L. F. R. Barrie, A. Hagborg, B. J. Crandall-Stotler, S. R. Gradstein, R. E. Stotler & M. von Konrat. 2015. Notes on early land plants today. 73. Genera of Lejeuneaceae established in the p
  6. Renzaglia, K. S., B. Crandall-Stotler, S. Pressel, J. G. Duckett, S. Schuette & P. K. Strother. 2015. Permanent spore dyads are not 'a thing of the past': on their occurrence in the liverwort Haplomitrium (Haplomitriopsida). Botanical Journal of the Linnean Society 179: 658-669.
  7. Villarreal A., J. C., B. J. Crandall-Stotler, M. L. Hart, D. G. Long & L. L. Forrest. 2015. Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiophyta) with a slow molecular rate. New Phytologist 209: 1734-1746.



  1. Gradstein, S. R., B. Laenen, J.-P. Frahm, U. Schwarz, B. J. Crandall-Stotler, J. J. Engel, M. von Konrat, R. E. Stotler, B. Shaw & A. J. Shaw. 2014. Taxonomic status of the enigmatic Phycolepidoziaceae (Marchantiophyta: Jungermanniales) with description of a new species, Phycolepidozia indica. Taxon 63: 498-508.

  2. Li, F.-W. et al. 2014. Horizontal transfer of an adaptive chimeric photoreceptor from bryophytes to ferns. PNAS 111: 6672-6677. [B. Crandall-Stotler is one of 35 authors on this paper; she contributed cultures of several of the hornworts used in the study.]

  3. Laenen, B. et al. 2014. Extant diversity of bryophytes emerged from successive post-Mesozoic diversification bursts. Nature Communications 5: Article # 5134. Published 27 October 2014. (doi:10.1038/ncomms6134) [B. Crandall-Stotler is one of the authors on this publication that is based on the results of the Liverwort Tree of Life dataset.]


      1. Stotler, R. E., B. Crandall-Stotler & N. Wickett. 2013. 2.2 Liverworts: Aneuraceae. pp. 20-23. IN: V. S. F. T. Merckx (ED.), Mycoheterotophy. The Biology of Plants Living on Fungi. Springer, Berlin. To access: Go to the Springer ad for the book Mycoheterotrophy Click on the bottom entry of that page "Download Sample pages 1 (pdf, s.9 MB)" and it will take you to Chapter 2 Taxonomy and Classification that includes the Liverworts entry.
      2. Söderström, L., B. Crandall-Stotler, R. E. Stotler, J. Váña, A. Hagborg & M von Konrat. 2013. Notes on Early Land Plants Today. 36. Generic treatment of Lophocoleaceae (Marchantiophyta). Phytotaxa 97: 36-43. (1 May 2013)
      3. Crandall-Stotler, B., R. E. Stotler, V. A. Bakalin & W. T. Doyle. 2013. A New Species of Mesoptychia (Lindb.) A. Evans from California. Polish Journal of Botany 58: 81-89.
      4. Söderström, L., J. Váña, B. Crandall-Stotler, R. E. Stotler, A. Hagborg & M von Konrat. 2013. Notes on Early Land Plants Today. 43. New combinations in Lophocoleaceae (Marchantiophyta). Phytotaxa 112 (1): 18–32. (17 June 2013)


      1. Crandall-Stotler, B. & R. E. Stotler. 2012. Ontogenetic studies, sporophyte anatomy and phylogenetic affinities of Phyllothallia nivicola (Phyllothalliaceae, Marchantiophyta). Nova Hedwigia 95: 277-294.


      1. Crandall-Stotler, B., R. E. Stotler, L. Zhang & L. L. Forrest. 2010. On the morphology, systematics and phylogeny of Noteroclada (Noterocladaceae, Marchantiophyta). Nova Hedwigia 91: 421450


      1. Crandall-Stotler, B, R. E. Stotler & D. G. Long. 2009. Phylogeny and classification of the Marchantiophyta. Edinburgh Journal of Botany 66: 155-198. [Copyright by Cambridge University Press <>]
      2. Bartholomew-Began, S. E. 2009. Juvenile gametophyte development in the Blasiales. 3. Sporeling ontogeny of Cavicularia densa. Bryologist 112: 354-358 [Copyright by The American Bryological and Lichenological Society, Inc.]


      1. Stotler, R. E. & B. J. Crandall-Stotler. 2008. Correct author citations for some upper rank names of liverworts (Marchantiophyta). Taxon 57: 289-292
      2. Crandall-Stotler, B, R. E. Stotler & D. G. Long. 2009[2008]. Morphology and classification of the Marchantiophyta, pp. 1-54. In B. Goffinet & A. J. Shaw (eds.), Bryophyte Biology, ed. 2. Cambridge University Press, Cambridge.


      1. Crandall-Stotler, B. J. & R. E. Stotler. 2007. On the identity of Moerckia hibernica (Hook.) Gottsche (Moerckiaceae fam. nov., Marchantiophyta). Nova Hedwigia, Beiheft 131: 41-59.


      1. Doyle, W. T. and R. E. Stotler. 2006. Contributions toward a bryoflora of California III. Keys and annotated species catalogue for liverworts and hornworts. Madroño 53: 89-197.
      2. Crandall-Stotler, B. J. & R. E. Stotler. 2006. Anatomical studies of Xenothallus vulcanicola R. M. Schust., a rare endemic liverwort from New Zealand. Journal of the Hattori Botanical Laboratory 100: 53-67.
      3. Forrest, L. L., E. C. Davis, D. G. Long, B. J. Crandall-Stotler, A. Clark and M. L. Hollingsworth. 2006. Unraveling the evolutionary history of the liverworts (Marchantiophyta): multiple taxa, genomes and analysis" by. Bryologist 109: 303-334.
      4. Qiu, Y.-L., L. Li, B. Wang, Z. Chen, V. Knoop, M. Groth-Malonek, O. Dombrovska, J. Lee, L. Kent, J. Rest, G. F. Estabrook, T. A. Hendry, D. W. Taylor, C. M. Testa, M. Ambros, B. Crandall-Stotler, R. J. Duff, M. Stech, W. Frey, D. Quandt, and C. C. Davis. 2006. The deepest divergences in land plants inferred from phylogenomic evidence. Proceedings of the National Academy of Sciences of the USA 103: 15511-15516