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Bryophyte Images
Monographic Studies
Phylogenetic Studies
Axenic Cultures

Bryophytes
Mosses, Liverworts & Hornworts

"Bryophytes" is a resource devoted to Bryology, the branch of plant science concerned with the study of mosses, liverworts and hornworts. It provides information on the classification, structural features, natural history, ecology and evolutionary relationships of these plants. Although small in stature, bryophytes play significant roles in diverse terrestrial ecosystems. They are found growing on soil, rocks and/or trees throughout the world, from coastal Antarctica to the peat bogs of the Northern hemisphere, from the deserts of Australia to the rain forests of the Amazon. They are an essential part of this planet's biodiversity. Click on any of the topics listed to learn more about these fascinating organisms.

Latest Site Features

April, 2015

  • Shaw, B., B. Crandall-Stotler, J. Vána, R. E. Stotler, M. von Konrat, J. J. Engel, E. C. Davis, D. G. Long, P. Sova & A. J. Shaw. 2015. Phylogenetic relationships and morphological evolution in a major clade of leafy liverworts (Phylum Marchantiophyta, Order Jungermanniales): Suborder Jungermanniineae. Systematic Botany 40: 27-45.

  • Söderström, L., J. Vána, B. Crandall-Stotler, M. A. M. Renner, A. Hagborg & M. von Konrat. 2015. Notes on early land plants today. 68. Miscellaneous notes on Marchantiophyta. Phytotaxa 202: 69-72.


  • January, 2015

  • Vitt, D. H. & B. Crandall-Stotler. 2014. Raymond Eugene Stotler (1940–2013) - Remembering an American hepaticologist. The Bryologist 117: 405-408.

  • Laenen, B. et al. 2014. Extant diversity of bryophytes emerged from successive post-Mesozoic diversification bursts. Nature Communications 5: Article # 5134. Published 27 October 2014.  (doi:10.1038/ncomms6134)  [B. Crandall-Stotler is one of the authors on this publication that is based on the results of the Liverwort Tree of Life dataset.]