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Bryophyte Images
Monographic Studies
Phylogenetic Studies
Axenic Cultures

Bryophytes
Mosses, Liverworts & Hornworts

"Bryophytes" is a resource devoted to Bryology, the branch of plant science concerned with the study of mosses, liverworts and hornworts. It provides information on the classification, structural features, natural history, ecology and evolutionary relationships of these plants. Although small in stature, bryophytes play significant roles in diverse terrestrial ecosystems. They are found growing on soil, rocks and/or trees throughout the world, from coastal Antarctica to the peat bogs of the Northern hemisphere, from the deserts of Australia to the rain forests of the Amazon. They are an essential part of this planet's biodiversity. Click on any of the topics listed to learn more about these fascinating organisms.

Latest Site Features

November, 2015

  • Renzaglia, K. S., B. Crandall-Stotler, S. Pressel, J. G. Duckett, S. Schuette & P. K. Strother. 2015. Permanent spore dyads are not 'a thing of the past': on their occurrence in the liverwort Haplomitrium (Haplomitriopsida). Botanical Journal of the Linnean Society 179: 658-669.

  • Villarreal A., J. C., B. J. Crandall-Stotler, M. L. Hart, D. G. Long & L. L. Forrest. 2015. Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiophyta) with a slow molecular rate. New Phytologist 209: 1734-1746.


  • April, 2016

  • Long, D. G., Forrest, L. L., Villarreal, J. C. & Crandall-Stotler, B. J. 2016. Taxonomic changes in Marchantiaceae, Corsiniaceae and Cleveaceae (Marchantiidae, Marchantiophyta). Phytotaxa 252: 77-80.

  • Söderström, L., Hagborg, A. , von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E. , Cargill, D. C., Costa, D. P., Crandall-Stotler, B. J. & 31 additional authors. 2016. World checklist of hornworts and liverworts. PhytoKeys 59: 1-828. (doi: 10.3897/phytokeys.59.6261). [This list of all recognized taxa of hornworts and liverworts was compiled by the first 3 authors, from contributions provided by the other authors involved in this very large project. All authors further edited and reviewed the manuscript as it was developed.]